Gully erosion is a major driver of land degradation globally, particularly in semi-arid regions where it is fundamentally controlled by rainfall and runoff dynamics. Understanding how rainfall translates into runoff in gullied landscapes is crucial for predicting erosion processes and modelling runoff to inform land management strategies. In this study, rainfall-runoff analysis was conducted using high-resolution rainfall and runoff data from intensely monitored alluvial gullies in the semi-arid regions of northern Australia. Runoff responses were strongly seasonal, with flashy but low-volume flows during the early wet season (October-November) and prolonged, high-discharge events during peak rainfall months (December-March). Antecedent soil moisture had a limited influence on runoff generation, likely due to rapid wetting-drying cycles and shallow infiltration depths. Notably, rainfall-runoff behavior diverged with catchment-to-gully area ratio (A(ca)): linear runoff to rainfall responses were observed where gullies were eroded to the catchment limit (A(ca) approximate to 1) whereas high-A(ca) systems (A(ca) > 5) exhibited threshold, stepwise behavior with upslope contributions activating at similar to 26 mm event rainfall. Field infiltration tests showed upslope catchment infiltration capacity was similar to 70% higher than on gully floors (similar to 36 vs. 21 mm h(-1)). This indicates greater near-surface storage and delayed upslope runoff, consistent with an activation threshold for upslope contributions. Mean rainfall-runoff ratios were higher in low-A(ca) gullies (approximate to 0.52-0.68) than in high-A(ca) systems (approximate to 0.40-0.46). These findings have implications for rainfall-runoff modelling, process-based understanding of gully erosion and gully management in semi-arid environments.