New insects from the Permian of Catalonia (Spain) predate the early diversification of Triassic clades adapted to arid environments (Grylloblattodea, Probnidae, Dictyoptera)

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  • Probnis sauvanyaensis sp. nov., the first representative of the 'grylloblattodean' family Probnidae from the Iberian Peninsula, is described from the middle to ?upper Permian outcrop of Sauvany & agrave; in the Catalan Pyrenees. The family is otherwise represented by two additional species of Probnis from the ?middle Permian of the USA and by a single genus and species from the ?Lower Triassic of Ukraine. As is known from outcrops with rather extreme climates, the family may have survived the global warming of the Permian-Triassic crisis. We also describe a roachoid of the family Spiloblattidae from the same outcrop as the second representative of this family from the Iberian Peninsula.