The semi-arid region of Brazil has broad phytophysiognomic heterogeneity. The forest formations in these regions comprise the upland forests, or `Brejos Nordestinos; which are fragments of humid forests surrounded by caatinga vegetation, a predominant vegetation type in the Brazilian semiarid region (Tabarelli & Santos 2004). These areas are refuges for organisms within the semi-arid domain and are associated with the occurrence of plateaus that confer mild climatic conditions and higher vegetation cover to the region, compared to the surrounded semi-arid ecosystems (Medeiros & Cestaro 2019).